In males, penis length is 9.6 to 11.8 mm ( 3⁄8 to 15⁄32 in) and baculum length is 8.8 to 10.0 mm ( 11⁄32 to 13⁄32 in). Copulation by male greater short-nosed fruit bats is dorsoventral and the females lick the shaft or the base of the male's penis, but not the glans which has already penetrated the vagina.
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Journal of Molluscan Studies. Most marsupials, except for the two largest species of kangaroos and marsupial moles  assuming the latter are true marsupials , have a bifurcated penis , separated into two columns, so that the penis has two ends corresponding to the females' two vaginas. Walther Includes Address 2 Phone 1. Stackpole Books. Tapirs have exceptionally long penises relative to their body size. Vertebrate Biology. Resides in Fenton, MO. Herrera 15 August During its waking hours, it remains hidden beneath a skin receptacle, until it becomes erect and it projects outside in a rostral direction. As of [update] , a systematic review of 15, men, and the best research to date on the topic, as the subjects were measured by health professionals, rather than self-measured, has concluded that the average length of an erect human penis is Even within the Vertebrata there are morphological variants with specific terminology, such as hemipenes. The papilla nipple-like projection on the dorsal upper side of the penis is covered with small spines, a character the marsh rice rat shares only with Oligoryzomys and Oryzomys couesi among oryzomyines examined. It is a passage both for urine and for the ejaculation of semen. External male genitalia of Papio hamadryas and Chlorocebus pygerythrus. Histological study of the cloacal region and associated structures in the hedgehog tenrec Echinops telfairi. Indiana University Press. Jackson, Hartley H. In Transandinomys talamancae , the outer surface of the penis is mostly covered by small spines, but there is a broad band of nonspinous tissue. Retrieved 15 July Biology, Medicine, and Surgery of Elephants. San Jose Mercury News. Retrieved 19 July Zeveloff Includes Address 5 Phone 2. Retrieved 5 May Archived from the original PDF on The Society. Resides in Bluefield, WV. Caro 15 August Wikimedia Commons has media related to Human penis. Discovery Publishing House. In del Hoyo, J. Retrieved 11 June Journal of the Mammalogical Society of Japan. Folia Primatol. Oslo, Norway: Meddelelser. Retrieved 28 June England, Gary 15 April In most species of animals in which there is an organ that might reasonably be described as a penis, it has no major function other than intromission, or at least conveying the sperm to the female, but in the placental mammals the penis bears the distal part of the urethra , which discharges both urine during urination and semen during copulation. Resides in Coxs Creek, KY. In Thomasomys ucucha the glans penis is rounded, short, and small and is superficially divided into left and right halves by a trough at the top and a ridge at the bottom. Frandson, Rowen D. Includes Address 2 Phone 2 Email 2. Barcelona, Spain: Lynx Edicions. Hoffmeister, Donald F. Statistics for all 73 Jamie Reece results:. Copulation by male greater short-nosed fruit bats is dorsoventral and the females lick the shaft or the base of the male's penis, but not the glans which has already penetrated the vagina.
The term penis applies to many intromittent organs , but not to all. As an example, the intromittent organ of most cephalopoda is the hectocotylus , a specialized arm, and male spiders use their pedipalps. Even within the Vertebrata there are morphological variants with specific terminology, such as hemipenes. In most species of animals in which there is an organ that might reasonably be described as a penis, it has no major function other than intromission, or at least conveying the sperm to the female, but in the placental mammals the penis bears the distal part of the urethra , which discharges both urine during urination and semen during copulation. Most male birds e. Among bird species with a penis are paleognathes tinamous and ratites  and Anatidae ducks, geese and swans. The lake duck also called Argentine blue-bill has the largest penis in relation to body size of all vertebrates; while usually about half the body size 20 cm , a specimen with a penis While most male birds have no external genitalia, male waterfowl Anatidae have a phallus. Most birds mate with the males balancing on top of the females and touching cloacas in a "cloacal kiss"; this makes forceful insemination very difficult. The phallus that male waterfowl have evolved everts out of their bodies in a clockwise coil and aids in inseminating females without their cooperation. These structures make it harder for males to achieve intromission. The clockwise coils are significant because the male phallus everts out of their body in a counter-clockwise spiral; therefore, a clockwise vaginal structure would impede forceful copulation. Studies have shown that the longer a male's phallus is, the more elaborate the vaginal structures were. The lake duck is notable for possessing, in relation to body length, the longest penis of all vertebrates ; the penis, which is typically coiled up in flaccid state, can reach about the same length as the animal himself when fully erect, but is more commonly about half the bird's length. The lake duck has a corkscrew shaped penis. Male and female emus are similar in appearance,  although the male's penis can become visible when it defecates. The male tinamou has a corkscrew shaped penis, similar to those of the ratites and to the hemipenis of some reptiles. Females have a small phallic organ in the cloaca which becomes larger during the breeding season. As with any other bodily attribute, the length and girth of the penis can be highly variable between mammals of different species. A bone called the baculum or os penis is present in most mammals but absent in humans, cattle and horses. In mammals the penis is divided into three parts: . The internal structures of the penis consist mainly of cavernous, erectile tissue , which is a collection of blood sinusoids separated by sheets of connective tissue trabeculae. Some mammals have a lot of erectile tissue relative to connective tissue, for example horses. Because of this a horse's penis can enlarge more than a bull's penis. The urethra is on the ventral side of the body of the penis. As a general rule, a mammal's penis is proportional to its body size, but this varies greatly between species — even between closely related ones. For example, an adult gorilla 's erect penis is about 4. In comparison, the human penis is larger than that of any other primate , both in proportion to body size and in absolute terms. The penises of even-toed ungulates are curved in an S-shape when not erect. When mating, the tip of a male pronghorn 's penis is often the first part to touch the female pronghorn. The penis of a dromedary camel is covered by a triangular penile sheath opening backwards,  and is about 60 cm 24 in long. Copulation time ranges from 7 to 35 minutes, averaging 11—15 minutes. Bulls have a fibro-elastic penis. Given the small amount of erectile tissue, there is little enlargement after erection. The penis is quite rigid when non-erect, and becomes even more rigid during erection. Protrusion is not affected much by erection, but more by relaxation of the retractor penis muscle and straightening of the sigmoid flexure. The male genitalia of mouse deer are similar to those of pigs. The purpose of this is not fully understood. A stag's penis forms an S-shaped curve when it is not erect, and is retracted into its sheath by the retractor penis muscle. Cetaceans' reproductive organs are located inside the body. Male cetaceans whales, dolphins, and porpoises have two slits, the genital groove concealing the penis and one further behind for the anus. The penis on a right whale can be up to 2. Stallions male horses have a vascular penis. When non-erect, it is quite flaccid and contained within the prepuce foreskin, or sheath. Tapirs have exceptionally long penises relative to their body size. All members of Carnivora except hyenas have a baculum. During copulation, the spotted hyena inserts his penis through the female's pseudo-penis instead of directly through the vagina , which is blocked by the false scrotum and testes. Once the female retracts her clitoris, the male enters the female by sliding beneath her, an operation facilitated by the penis's upward angle. Domestic cats have barbed penises, with about — one millimeter long backwards-pointing spines.